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After searching the the Banished stronghold that Jerome-092 and the UNSC destroyed before fighting Decimus, they find intel informing them that the banished are utilizing the power of The Cartographer on the Ark, which is allowing them to find key systems and weapons on the Ark. Due to the strong Forerunner defenses, the only way the UNSC can get into and use The Cartographer is to capture the surrounding forerunner defense towers so that Anders can gain control of the elevator and take the UNSC up and inside.
Jerome is on the ground once again and is accompanied by a small ground force including Warthogs and Cyclops'. They traverse the land to a designated base building site, encountering small groups of Banished light and heavy vehicles, one of which being a Wraith that Jerome hijacks. After the base is established, Jerome alongside a decent sized UNSC force of infantry and air vehicles take and maintain control of the defense towers long enough for Anders to gain control of the elevator to The Cartographer, all the way fighting off a large Banished ground force as well as destroying some smaller Banished structures.